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Plant Physiological Ecology,Used
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Box 9E. 1 Continued FIGURE 2. The CSR triangle model (Grime 1979). The strategies at the three corners are C, competiti winning species; S, stresstolerating s cies; R,ruderalspecies. Particular species can engage in any mixture of these three primary strategies, and the m ture is described by their position within the triangle. comment briefly on some other dimensions that Grimes (1977) triangle (Fig. 2) (see also Sects. 6. 1 are not yet so well understood. and 6. 3 of Chapter 7 on growth and allocation) is a twodimensional scheme. A CS axis (Com titionwinning species to Stresstolerating spe Leaf Economics Spectrum cies) reflects adaptation to favorable vs. unfavorable sites for plant growth, and an R Five traits that are coordinated across species are axis (Ruderal species) reflects adaptation to leaf mass per area (LMA), leaf lifespan, leaf N disturbance. concentration, and potential photosynthesis and dark respiration on a mass basis. In the fivetrait TraitDimensions space,79%ofallvariation worldwideliesalonga single main axis (Fig. 33 of Chapter 2A on photo A recent trend in plant strategy thinking has synthesis; Wright et al. 2004). Species with low been traitdimensions, that is, spectra of varia LMA tend to have short leaf lifespans, high leaf tion with respect to measurable traits. Compared nutrient concentrations, and high potential rates of massbased photosynthesis. These species with category schemes, such as Raunkiaers, trait occur at the quickreturn end of the leaf e dimensions have the merit of capturing cont nomics spectrum.
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